In addition, most firefly species have been regarded as habitat and/or season specialists as adults ( Silveira et al., 2020). Because signal diversity is associated with sensor morphology, the use of light and pheromones is accompanied by enlarged eyes and complex antennal morphology, respectively ( Stanger-Hall et al., 2018). For example, adults may use diverse mating signals, including light-as continuous glows or flashes-and pheromones, alone or in combination ( De Cock & Matthysen, 2005 Lewis & Cratsley, 2008 Stanger-Hall et al., 2018). The charismatic fireflies (Coleoptera: Lampyridae) are famous for their flashing adults, but the nearly 2200 described species worldwide show wide-ranging phenotypic and ecological diversity, including the less appreciated diurnal and dark species ( Branham, 2010 Martin et al., 2019).
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